Name:
Nanotyrannus
(Dwarf tyrant).
Phonetic: Nah-no-ty-ran-nus.
Named By: Robert T. Bakker, Phil Currie
& Michael Williams - 1988.
Synonyms: Gorgosaurus lancensis, Deinodon
lancensis, Aublysodon lancensis, Albertosaurus lancensis.
Classification: Chordata, Reptilia, Dinosauria,
Saurischia, Theropoda, Tyrannosauridae.
Species: N. lancensis (type).
Diet: Carnivore.
Size: Approximately 5.2 meters long for holotype.
Known locations: North America.
Time period: Maastrichtian of the Cretaceous.
Fossil representation: Single skull.
Nanotyrannus
is probably the most controversial member of the Tyrannosauridae
because while some palaeontologists think that it represents a distinct
genus, others think that it is simply a juvenile form of a larger
tyrannosaurid genus, or even a dwarf species of another genus.
Indeed when the skull was first described by Charles W. Gilmore in
1946 it was classified as a species of Gorgosaurus.
The
controversy surrounding Nanotyrannus began in
1988 when during a
re-examination of the skull by Robert T. Bakker, Phil Currie and
Michael Williams, the skull bones where perceived to have been fused
together. This is typically a sign of an adult individual as the
skull bones of individual animals remain separate while in the juvenile
stages for easier growth. When the growth begins to slow down the
bones become fused together for additional strength during the animal’s
adult life.
However
in 1999 another palaeontologist named Thomas Carr conducted a
detailed analysis on the skull and found that the bones were not
actually fused together after all. This observation split
palaeontologists all over the world into thinking that Nanotyrannus
was
either a small tyrannosaurid or a juvenile of a larger one. Debate
continued to sway one way or the other until the discovery of a new
tyrannosaur specimen that was nicknamed ‘Jane’. Not only was Jane
the same size and species as Nanotyrannus, but
Jane was without doubt
a juvenile tyrannosaur. In light of this new discovery many of the
palaeontologists who had supported the idea that Nanotyrannus
was a new
genus now switched to the idea that Nanotyrannus
really was a juvenile
of another genus, including Phil Currie and Michael Williams.
Other palaeontologists still however think that Nanotyrannus
deserves to be placed within its own genera because it is still
different enough from other known forms.
Initially
Nanotyrannus was thought to be a juvenile
tyrannosaurid (possibly of
Tyrannosaurus)
because of the large number of teeth that were present
a study that showed a correlation between tyrannosaurids reducing the
numbers of teeth as they aged. The problem with this study however is
that it only supplied a few examples, and not only do some genera
like Tarbosaurus show roughly the same amount of
teeth between adults
and juveniles, but other genera show a great discrepancy between the
number of teeth present in individuals of the same age and genus.
This is why teeth count is no longer considered a reliable identifying
feature for age and genus.
There
only remains one feature that is present upon the Nanotyrannus
holotype
and the Jane specimen that may save Nanotyrannus as
being a valid genus
and this is a small pit in the quadratojugal at the back of the skull.
This remains unknown in the specimens of other juvenile tyrannosaurids
and is so far not present in any known adults. Unless this proves to
be a feature that grew out as the animal reached adulthood it may
become the identifying feature of the genus.