Name: Voay.
Phonetic: Vo-ay.
Named By: C. A. Brochu - 2007.
Synonyms: Crocodylus robustus.
Classification: Chordata, Reptilia,
Crocodylidae, Crocodylinae.
Species: V. robustus (type).
Diet: Carnivore.
Size: Estimated up to 5 meters long.
Known locations: Madagascar.
Time period: Holocene.
Fossil representation: Skulls, vertebrae,
osteoderms.
The
remains of the crocodile
Voay were first described
as a species of
Crocodylus (C. robustus)
back in 1872 by Grandidier and
Vaillant, however later study found that the remains were nearer to
the dwarf crocodile Osteolaemus, although not
close enough to be
included with this genus either. For this reason C. A. Brochu
created a new genus under the name Voay in 2007,
and in keeping
with standard practices for naming new genera and species from existing
classified material, the species name of robustus
was retained to
create a new type species of Voay robustus.
The
remains of Voay which include skulls, vertebrae
and osteoderms (the
bony armour plates in the skin that are also sometimes called
‘scutes’) are classed as sub fossils which mean that they have not
been completely fossilised. These remains are also dated to as
recently as two thousand years ago, something that suggests that the
genus went extinct with the arrival of the first humans upon
Madagascar. This also coincides with the disappearance of other large
animals from Madagascar at this time which has led to theory that human
hunting may have been a factor in the disappearance of these
animals. Although Voay is dated during the
Holocene, a
disappearance two thousand years ago would on paper suggest that the
genus only lived ten thousand years. Realistically however, Voay
probably also existed into the at least the late Pleistocene era, but
further fossil remains from different time periods would be needed to
establish and confirm a more complete temporal range for this genus.
The
Nile crocodile (Crocodylus niloticus) is also
known from
Madagascar, but there is some debate concerning whether the Nile
crocodile and Voay co-existed in Madagascar at the
same time, or if
Nile crocodiles colonised Madagascar to fill a predatory void left
behind by the disappearance of Voay. At this
point either theory is
plausible since although similarly sized, Voay is
morphologically
different to the C. niloticus. One key area is
the more robust
limbs, and since crocodiles usually rely upon their tails for
swimming, the stronger limbs might suggest that Voay
spent a greater
amount of time on the land than C. niloticus.
This could also mean
a stronger disposition towards hunting and possibly even scavenging on
land, which would infer different specialisation and behaviour to the
C. niloticus.
Voay
also has a proportionately shorter and deeper snout than C.
niloticus
with nasal openings that are also reduced in size. This also infers a
different prey specialisation to the longer and more slender snouted
C. niloticus and if the two lived in the same
habitats, it’s
possible that they may have been able to coexist by adopting different
lifestyles. A precedent could be established by looking at South
American crocodiles of the Miocene. Here you had three giant
crocodiles called Purussaurus,
Gryposuchus
and Mourasuchus.
All
three lived in the same approximate geographic area but each
specialised in a different feeding strategy. Purussaurus
had a skull
similar to a caiman, indicating that it was a hunter of large prey.
Gryposuchus however was more like a gharial
(alternatively gavial)
and was suited to hunting fish. Finally Mourasuchus
seems to have
been a filter feeder that consumed large amounts of small prey
animals. Even though we still don’t know exactly how well these three
crocodiles got on, it is possible that they could inhabit the same
water systems without competing for the same food as one another.
Voay
did have a pair of horns that grew from the back of its head that were
actually extensions of the squamosal bone (one of the most rearward
portions of the skull). These horns are often seen in other
crocodiles as well, which has led to the more common term of these
species and genera being called ‘horned crocodiles’.
Further reading
- Description of a skull of the extinct Madagascar crocodile, Crocodilus
robustus Vaillant and Grandidier. - Bulletin of the American
Museum of
Natural History 44 (4): 25. - Charles C. Mook.
- The giant dwarf crocodile: a reappraisal of ‘Crocodylus’
robustus
from the Quaternary of Madagascar. In: Patterson, Goodman &
Sedlock, eds., Environmental Change in Madagascar. p. 70. - C. A.
Brochu & G. W. Storrs - 1995.
- Morphology, relationships, and biogeographical significance of an
extinct horned crocodile (Crocodylia, Crocodylidae) from the Quaternary
of Madagascar. - Zoological Journal of the Linnean Society 150:835-863.
- C. A. Brochu - 2007.
- The late Pleistocene horned crocodile Voay robustus
(Grandidier
& Vaillant, 1872) from Madagascar in the Museum f�r Naturkunde
Berlin. - Fossil Record 12: 13. - C. Bicklemann, N. Klein - 2009.