Name:
Pliohippus
(Pliocene horse/More horse - refer to main text).
Phonetic: Ply-oh-hip-pus.
Named By: Othniel Charles Marsh - 1874.
Synonyms: Dinohippus subvenus, Hippidium
robustus, Merychippus campestris, Pliohippus lullianus, Pliohippus
robustus, Protohippus robustus.
Classification: Chordata, Mammalia,
Perissodactyla, Equidae.
Species: P. castilli, P. fossulatus,
P. mirabilis, P. nobilis, P. pernix, P. tantalus, P.
tehonensis. *Note - Other species may be missing.
Diet: Herbivore.
Size: About 1.2 meters tall at the shoulder,
depending upon species.
Known locations: Canada and USA.
Time period: Messinian of the Miocene through to the
early Pleistocene.
Fossil representation: Multiple specimens.
In
the on-going evolution of horses Pliohippus
represents the next stage
after forms like Merychippus.
The most noted feature of Pliohippus
is
that it has even more developed hoof feet supported by the middle toe,
with the two side toes being reduced so much that in life they would
have been barely visible if at all. It is still under debate however
as to how close Pliohippus was to modern horses
that come under the
Equus genus. Pliohippus was
certainly ancestral
but first may have
given rise to other forms such as Astrohippus
or Dinohippus
which are
turn are thought to be more closely related to modern forms due to
their even greater similarity.
The
key features that differentiate Pliohippus from
modern forms are the
presence of two pits in the skull that are in front of the eyes, a
feature unknown in modern forms. Explanations for these pits have
included space to accommodate larger face muscles to even resonating
chambers that allowed Pliohippus to adjust the
sounds of its calls.
Despite these theories however, the function of these pits remains
largely unknown.
Pliohippus
is often credited with meaning ‘Pliocene horse’ because it was once
thought to live during the Pliocene (the boundary between the Miocene
and Pliocene has been re-established since Pliohippus
was named),
however the ‘Plio’ part actually translates as ‘more’.
However in this frame of reference Pliohippus can
still mean ‘more
like a horse than other (earlier) prehistoric horses’.
Further reading
- Notice of new equine mammals from the Tertiary formation. - American
Journal of Science 7(39):247-258. - O. C. Marsh - 1874.
- Equidae from the Pliocene of Texas. University of California
Publications. - Bulletin of the Department of Geological Sciences
19(17):349-396. - W. D. Matthew & R. A. Stirton - 1930.
- On the Equid genus Dinohippus Quinn 1955 and
Pliohippus Marsh 1874. -
Bolletino della Societa Paleontologica Italiana 27(1):61-72. - A.
Azzaroli - 1988.
- The evolution of Oligocene horses. In D. R. Prothero and R. M. Schoch
(eds.) - The Evolution of Perissodactyls 142-175. - D. R. Prothero
& N. Shubin - 1989.